The International Lepidoptera Survey (TILS-TTR)

Main | News | Taxonomic Report | Type Photos | Lep Names | Online Library | Links | Leps-Talk | About

The International Lepidoptera Survey - The Taxonomic Report

Volume 1 | Volume 2 | Volume 3 | Volume 4 | Volume 5 | Volume 6 | Volume 7 | Volume 8 | Volume 9 | Volume 10 | Volume 11

Volume
Dates
Contents
Volume 1
1998-1999

An Addendum to Anthocharis midea Dos Passos and Klots 1969. (Description of a new subspecies from Texas.)

The rediscovery, taxonomy, and biology of Chlosyne gorgone gorgone and Chlosyne ismeria (Nymphalidae) in Burke County, Georgia

Two new Nymphalidae from western North Carolina: new subspecies of Speyeria aphrodite and Phyciodes batesii

Subspecific status of southeastern U.S. Megathymus cofaqui and M. yuccae: renaming of the Florida subspecies of M. cofaqui

A comment on Friedlander’s Asterocampa (Nymphalidae, Apaturinae): designation of neotypes for A. celtis and A. clyton

An evolutionary subspecific assessment of Deciduphagus henrici (Lycaenidae) based on its utilization of Ilex and Non-Ilex hosts: description of a third Ilex associated subspecies. Designation of a neotype and type locality for Deciduphagus irus

A new subspecies of Brephidium isophthalma (Lycaenidae: Polyommatinae) from coastal South Carolina

Hübner’s helicta: the forgotten Neonympha. The recognition and elevation of Neonympha helicta (Nymphalidae: Satyrinae) to specific status. The designation of neotypes for N. helicta and N. areolatus. The subspecific transfer of septentrionalis to helicta and the description of a third helicta subspecies from south Florida

Celastrina idella (Lycaenidae: Polyommatinae): a new butterfly species from the Atlantic Coastal Plain

Three new Hesperioidae (Hesperiinae) from South Carolina: new subspecies of Euphyes bimacula, Poanes aaroni, and Hesperia attalus.

Volume 2
2000

A new elfin butterfly (Lycaenidae: Eumaeini) from northern China with comments on the nomenclature of Palaearctic Elfins

Description of a new subspecies of Poanes aaroni (Hesperioidae: Hesperiinae) from the west central Gulf coast of the southern United States

A new species of Angulopis (Lycaenidae, Eumaeini) from relict coastal forest in east-central Argentina

A new North American swallowtail butterfly: description of a relict subspecies of Pterourus troilus (Papilionidae) from the southern tip of Florida

The biology, life history, and taxonomy of Celastrina neglectamajor (Lycaenidae: Polyommatinae)

Records for the utilization of Prunus as a larval foodplant by 71 species of Lepidoptera in northeast California

Nine new species of Lacinipolia (Noctuidae) from Arizona, California and vicinity

Volume 3
2001-2002

Clarification of and comments on northern Speyeria hydaspe subspecies (Lepidoptera: Nymphalidae)

An examination of southeastern U.S. Satyrium (Lycaenidae: Theclinae).
Part one: an obscure new subspecies of Satyrium edwardsii

An examination of southeastern U.S. Satyrium (Lycaenidae: Theclinae).
Part two: the identification and delimitation of nominate Satyrium liparops and the description of a new subspecies from west central peninsular Florida

An examination of the Mitoura (Lycaenidae) in the southeastern United States: with the description of a new subspecies of Mitoura hesseli

Initial survey of the butterflies and skippers in the vicinity of the Buck Creek serpentine barrens, Clay County, North Carolina

Name-bearing types and taxonomic synopsis of three lycaenid butterfly taxa from western Canada (Lepidoptera: Lycaenidae)

Pterourus appalachiensis (Papilionidae: Papilioninae), a new swallowtail butterfly from the Appalachian region of the United States

A review of Poanes hobomok (Hesperiidae: Hesperiinae), with the description of a new subspecies from the southern Appalachians

Taxonomy and distribution of butterflies (Papilionoidea) of the Skardu region, Pakistan

Volume 4
2003-2004

Fixation of type locality for Lycaena acmon Westwood and characterization of the species and its distribution

Butterflies of the North Okanagan, British Columbia, Canada

A subspecific assessment of the genus Hesperia (Hesperiinae) in eastern North America (part I: the south) new subspecies of Hesperia metea, Hesperia sassacus and Hesperia meskei

A taxonomic review of Chlosyne ismeria with description of a new subspecies from the southern Appalachian mountains

A case for taxonomic recognition of the taxon Enodia anthedon borealis A.H. Clark (Satyridae)

A taxonomic examination of Harkenclenus titus (Lycaenidae: Theclinae) in the eastern United States

A concise update of the information provided in The Butterflies of Southern California (1973)

Description of a multilevel cryptic new species of Phyciodes (Nymphalidae: Melitaeinae) from the southern Appalachian mountains

Volume 5
2004-2007

The Yosemite butterflies — text and plates

Volume 6
2004-2007

Discovery of a black female form of Pterourus appalachiensis (Papilionidae: Papilioninae) and additional observations of the species in West Virginia

Immature stages of Colias occidentalis sullivani from Oregon (Lepidoptera: Pieridae)

Immature stages of Colias johanseni from Arctic Canada (Lepidoptera: Pieridae)

Immature stages of Estigmene acrea from Guatemala (Lepidoptera: Arctidae)

Taxonomic analysis of the genus Megisto (Satyridae) in the eastern United States
Part I: types, type localities, and taxonomic relationships of the avaliable names

Celastrina serotina (Lycaenidae: Polyommatinae): a new butterfly species from the northeastern United States and eastern Canada

A new subspecies of Colias gigantea from arctic Alaska (Pieridae)

Bionomic studies of Papilio demoleus Linnaeus, the Citrus Butterfly, (Lepidoptera: Papilionidae), from Lower Sindh, Pakistan

Volume 7
2007-2014

A description of a new subspecies of Lycaena phlaeas (Lycaenidae: Lycaeninae) from Montana, United States, with a comparative study of old and new world populations

Natural life histories of Alaska Colias (Lepidoptera: Pieridae)

Tips on collecting and rearing immatures of 375 butterfly and skipper taxa

Observations on Anthocharis julia browningi and Anthocharis thoosa thoosa including tension zones near Nephi, Juab County, Utah

The nomenclatural status of ten names in the genus Pieris (Lepidoptera: Pieridae)

Larval host plants of Enodia anthedon, Satyrodes appalachia and S. eurydice in Vermont, USA

A case of sympatric Celastrina ladon (Cramer), Celastrina lucia (W. Kirby) and Celastrina neglecta (Edwards) (Lycaenidae: Polyommatinae) in Northern Virginia, with additional records of C. lucia in Virginia

Cercyonis pegala agawamensis (Satyridae): A new butterfly subspecies from the coastal salt marshes of the northeastern United States of America

Volume 8
2019-2020

Expanded phenotypic diagnoses for 24 recently named new taxa of Hesperiidae (Lepidoptera) — Nick V. Grishin

Changes to North American butterfly names — Jing Zhang, Qian Cong, Jinhui Shen, Paul A. Opler and Nick V. Grishin

Designation of a Neotype for Oeneis melissa (Fabricius, 1775) and a Lectotype for Oeneis polixenes (Fabricius, 1775) (Lepidoptera: Nymphalidae: Satyrinae) — Crispin S. Guppy

Designation of neotype of Hemileuca maia (Drury, 1773) and refinement of its type locality (Bombycoidea, Saturniidae, Hemileucinae) — Harry Pavulaan

A phenotypic comparison of regional populations of Hemileuca maia (Drury, 1773) with designations of new subspecies (Bombycoidea, Saturniidae, Hemileucinae) — Harry Pavulaan

Re-evaluation of Euchloe olympia (W. H. Edwards, 1871) (Pieridae: Pierinae) from a geographic and taxonomic perspective — Harry Pavulaan

Genomic evidence suggests further changes of butterfly names — Jing Zhang, Qian Cong, Jinhui Shen, Paul A. Opler, and Nick V. Grishin

Review of the Speyeria egleis complex in Montana, with the description of two new subspecies (Lepidoptera: Nymphalidae: Heliconiinae) — Steve Kohler

Volume 9
2020-2021

Evaluation of the taxonomic status of Eurytides marcellus form "floridensis" (W. Holland, 1898) (Papilionoidea, Papilioninae, Leptocircini) — Harry Pavulaan

Pupal polymorphism of the Bog Copper butterfly Lycaena (Epidemia) epixanthe (Bsd. & Le C. [1835]) (Lycaenidae: Lycaeninae) in New Jersey — David M. Wright

Genomics-guided refinement of butterfly taxonomy — Jing Zhang, Qian Cong, Jinhui Shen, Paul A. Opler, and Nick V. Grishin

Review of Montana Euphilotes Mattoni, [1978], with descriptions of new taxa (Lepidoptera: Lycaenidae: Polyommatinae) — Steve Kohler and Andrew D. Warren

Notes on Eastern North American Butterflies — David J. Hoag, Harry Pavulaan, Brian Taber, Kenneth Lorenzen, Richard D. Ullrich

Egg Plastron of the Bog Copper butterfly Tharsalea (Epidemia) epixanthe (Bsd. & Le C. [1835]) (Lycaenidae: Lycaeninae) — David M. Wright

Two new species of Hermeuptychia from North America and three neotype designations (Nymphalidae: Satyrinae) — Qian Cong, Eduardo P. Barbosa, Mario A. Marín, André V. L. Freitas, Gerardo Lamas, and Nick V. Grishin

Type locality of Polyommatus lucia (W. Kirby, 1837) (Lycaenidae: Polyomatinae) with an inspection of the species’ phenotypic expression — Norbert G. Kondla, Harry Pavulaan, and David M. Wright

Subspecific designation of the U.S.A. Interior Highlands population of Argynnis (Speyeria) diana (Cramer, 1777) (Nymphalidae: Heliconiinae: Argynnini: Argynnina) — Harry Pavulaan

Reevaluation of the described subspecies of Euphydryas phaeton (Drury, 1773) with a replacement name for Melitaea phaeton schausi (Clark, 1927) — Harry Pavulaan

Volume 10
2022

Natural history of the Guatemalan copper Iophanus pyrrhias (Godman and Salvin, 1887) (Lepidoptera: Lycaenidae) in Guatemala — José Monzón-Sierra and Gregory R. Ballmer

Celastrina ladon and C. neglecta are distinct species differentiated primarily by a diagnostic wing scale structure, voltinism, and host tolerance of Cornus florida — Harry Pavulaan

A rare case of mosaic gynandromorphism in the Zabulon Skipper (Lon zabulon) — Annette Allor

Rhode Island, USA Fall Lepidoptera Survey 2021 — Harry Pavulaan

A new species of Celastrina from the northwestern United States and southwestern Canada with a lectotype designation of Lycaena pseudargiolus var. nigrescens Fletcher (Lycaenidae: Polyommatinae) — Caitlin C. LaBar, Jonathan P. Pelham, Norbert G. Kondla

Genomic DNA sequencing reveals two new North American species of Staphylus (Hesperiidae: Pyrginae: Carcharodini) — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, and Nick V. Grishin

Checking the checkered taxonomy of Plötz's checkered skippers (Hesperiidae: Pyrgini) — Jing Zhang, Qian Cong, John M. Burns, and Nick V. Grishin

Subspecific designation for the Central Appalachian Mountains population of Argynnis (Speyeria) atlantis (W. H. Edwards) — Harry Pavulaan

Taxonomic discoveries enabled by genomic analysis of butterflies — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, Riley J. Gott, Pierre Boyer, Crispin S. Guppy, Steve Kohler, Gerardo Lamas, Paul A. Opler, and Nick V. Grishin

Neotype designation for Papilio fulgerator Walch, 1775 (Hesperiidae: Eudaminae) — Jing Zhang, Qian Cong, Gerardo Lamas, and Nick V. Grishin

Life history and ecology of the San Emigdio blue butterfly (Lepidoptera: Lycaenidae) — Gregory R. Ballmer

Establishment of the invasive Cactus Moth, Cactoblastis cactorum (Berg) (Lepidoptera: Pyralidae) in Pakistan: a potential threat to cultivated, ornamental and wild Opuntia spp. (Cactaceae) — Muhammad Ather Rafi, Harry Pavulaan, Muhammad Islam, Muhammad Ashfaq, Haseeb Kamran, Walija Fayaz, Gul Naz Parveen, Riffat Sultana, Ahmad Zia, Waqar Ahmed, Qudrat Ullah, Muhammad Qasim, Falak Naz, Nazeer Ahmed, Muhammad Tariq Khan, Muhammad Saeed, Jalal Hayat Khan

Volume 11
2023-2024

Additional taxonomic refinements suggested by genomic analysis of butterflies — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, Paul A. Opler, and Nick V. Grishin

A taxonomic list of the Old World genera in the subfamily Hesperiinae (Hesperiidae) arranged into tribes — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, and Nick V. Grishin

Butterfly classification and species discovery using genomics — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, and Nick V. Grishin

Examination of the status of Phyciodes tharos distincta Bauer, 1975, confirming it as a valid subspecies — Harry Pavulaan

Reassessment of Amblyscirtes hegon (Hesperiidae) as a complex of four distinct species revealed by genomic analysis — Harry Pavulaan, Ricky Patterson, and Nick V. Grishin

Genomic analysis reveals new species and subspecies of butterflies — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, and Nick V. Grishin

Taxonomic advances driven by the genomic analysis of butterflies — Jing Zhang, Qian Cong, Jinhui Shen, Leina Song, and Nick V. Grishin

Rhode Island, USA Fall Butterfly Survey 2023 — Harry Pavulaan

A review of the status of Poanes massasoit hughi Clark, 1931, confirming status as a range-wide variant form of P. massasoit (Scudder, 1863) — Harry Pavulaan

An updated geographic analysis of regional variation in Anthocharis midea (Hübner, [1809]) and review of dos Passos & Klots (1969) and Gatrelle (1998) — Harry Pavulaan

Volume 12
2024-

Determination of a new spring-flying species of the Pterourus glaucus complex (Papilionidae) in southern New Englands — Harry Pavulaan

The seasonal forms of Heraclides cresphontes Cramer, 1777, and clarification of the status of H. c. pennsylvanicus F. H. Chermock & R. L. Chermock, 1945 — Harry Pavulaan

Distribution of Edith's Copper Tharsalea editha in Canada — Janice E. Arndt

Addendum to: Determination of a new spring-flying species of the Pterourus glaucus complex (Papilionidae) in southern New England — Harry Pavulaan